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!{Html.TextBox("registerDetails.OtherCompanyActivity", registerModel.EditForm.OtherCompanyActivity, new { @class = "further_info", @tabindex = "15" })}

this is my dropdownlist. And when i have selected a value in the dropdownlist , the textbox should be made readonly.

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Use JavaScript
add/bind OnChange event to the dropdown list 

OtherCompanyActivity", registerModel.EditForm.OtherCompanyActivity, new { @class = "further_info", @tabindex = "15", onchange="dpwnChnaged()" })}

Check the value change from default value and make the text box ReadOnly by adding ReadOnly attribute to text box

I would prefer JQuery

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